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The terrestrial Adephaga of Iowa (Part 1) by Fanny Chastina Thompson Wickham, 1895

The terrestrial Adephaga of Iowa by Fanny Chastina Thompson Wickham, 1895, Page 11

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consist each of four pieces separated by sutures. The four are known as proscutum, scutum, scutellum and postscutellum. The ventral surface consists of the same pieces as the prothorax; i.e., mesosternum with its episterna and epinera and metasternum with the same. Wings. The anterior or mesothoracic wings are horny plates called elytra. The vary greatly in size, shape and sculpture. They usually cover the dorsal surface of the abdomen, but in many genera they are much shorter; sometimes they are united so as to form a single piece, and sometimes, but rarely, they are wanting. The metathoracic or posterior wings are membranous but with few nerves, so arranged as to allow of the complete folding of the wings under the elytra. Frequently the wings are absent. Legs. The first joint, that by which they are attached
 
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